Michael Shermer: Premoral Sentiments In Social Animals.

Barbara King argues that while primates may not possess morality in the human sense, they do exhibit some traits that would have been necessary for the evolution of morality. These traits include high intelligence, a capacity for symbolic communication, a sense of social norms, realization of "self", and a concept of continuity. Frans de Waal and Barbara King both view human morality as having grown out of primate sociality.

Many social animals such as primates, dolphins and whales have shown to exhibit what Michael Shermer refers to as premoral sentiments. According to Shermer, the following characteristics are shared by humans and other social animals, particularly the great apes: 

attachment and bonding, cooperation and mutual aid, sympathy and empathy, direct and indirect reciprocity, altruism and reciprocal altruism, conflict resolution and peacemaking, deception and deception detection, community concern and caring about what others think about you, and awareness of and response to the social rules of the group.

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